Principal Engineer, AI Security

Lila Sciences

• $252K — $374K *
Information Technology
Less than 5 years of experience
Job Overview by Ladders

Qualifications

  • Strong background in security engineering, cloud security, or DevSecOps with hands-on experience in production infrastructure.
  • Working knowledge of AI/ML systems and their lifecycle, including AI-specific risks.
  • Experience in software supply chain security and vulnerability management.
  • Proven ability to collaborate across engineering teams for effective remediation.
  • Comfortable operating in a fast-paced, ambiguous research environment.

Responsibilities

  • Translate the AI security roadmap into actionable controls with measurable outcomes.
  • Design and implement security controls for AI training and inference pipelines.
  • Ensure continuous compliance of AI infrastructure with internal standards.
  • Drive closure on vulnerability remediation across AI systems and dependencies.
  • Reduce risk through threat modeling and adversarial assessments.
  • Collaborate with cross-functional teams to embed security in the design phase.

Benefits

  • Comprehensive medical, dental, and vision coverage.
  • Employer-paid life and disability insurance.
  • Flexible time off with generous company-wide holidays.
  • Paid parental leave and educational assistance program.
  • Commuter benefits, including bike share memberships.
  • Company-subsidized lunch program.
Full Job Description
Your Impact at LILA

This role sits on the IT & Security team and owns security engineering for Lila's AI and ML systems, from training and inference pipelines to model registries, MCP servers, and agent runtimes.

As the senior individual contributor for AI security, you will translate the AI security roadmap into delivered controls, drive vulnerability remediation to closure, and reduce risk across the full AI lifecycle: data, models, pipelines, and agentic systems. The work is hands on, with technical depth across cloud security, supply chain security, and AI-specific threat models.

The role is deeply cross-functional. You will partner with AI Platform, AI Research, ML Operations, Science, and AI Safety so security shows up at design time rather than after the fact. Success looks like AI infrastructure that stays continuously compliant, measurable reduction in open risk, and engineering teams that treat security as part of how they build.

What You'll Be Building
  • Turn the AI security roadmap into delivered controls with measurable outcomes
  • Design and implement security controls across training and inference pipelines, model registries, MCP servers, and agent runtimes
  • Keep AI infrastructure continuously compliant against internal standards and applicable frameworks
  • Drive vulnerability remediation across AI infrastructure, models, and dependencies to closure with system owners
  • Reduce risk across data, models, pipelines, and agentic systems through threat modeling, control design, and adversarial assessment
  • Partner with AI Platform, AI Research, ML Operations, Science, and AI Safety so security is embedded at design time

What You'll Need to Succeed
  • Strong background in security engineering, cloud security, or DevSecOps, with hands-on experience securing production infrastructure
  • Working knowledge of AI/ML systems and the ML lifecycle, plus AI-specific risks such as prompt injection, model and data poisoning, insecure agent tooling, and inference-time threats
  • Experience with software supply chain security and dependency and vulnerability management
  • Experience implementing data security controls (classification, data flow mapping, access control, encryption) in cloud environments
  • Proven ability to partner across engineering teams and drive cross-functional remediation to closure
  • Comfort operating with ambiguity in a fast-moving research environment

Bonus Points For
  • Hands-on AWS security experience
  • Experience with SBOM tooling at scale
  • Familiarity with AI governance and assurance frameworks such as NIST AI RMF
  • Experience with MAESTRO-style threat modeling for agentic systems


Compensation

We offer competitive base compensation with bonus potential and generous early-stage equity. Your final offer will reflect your background, expertise, and expected impact.

U.S. Benefits. Full-time U.S. employees receive a comprehensive benefits program including medical, dental, and vision coverage; employer-paid life and disability insurance; flexible time off with generous company wide holidays; paid parental leave; an educational assistance program; commuter benefits, including bike share memberships for office based employees; and a company subsidized lunch program.

International Benefits. Full-time employees outside the U.S. receive a comprehensive benefits program tailored to their region. USD salary ranges apply only to U.S.-based positions; international salaries are set to local market.

Expected Base Salary Range

$252,000-$374,000 USD

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